Mystacocarida, Ostracoda (seed shrimps, mussel shrimps, Copepoda (cyclops), Remepedia, Cephalocardia, and Tantulocardia

Class Mystacocarida

This Class was first described in 1943. The first specimens were collected from beeches in Massachusetts and Connecticut. There are 12 species described so far; all are marine. They are found in interstitial spaces in shallow sediments. All are less than 0.5 mm long and show many primitive characteristics, see right.

Class Tantulocardia

There are twelve species, all microscopic 0.15 - 0.20 mm, ectoparasites of other marine crustaceans. The larvae and females attach to the host by an adhesive disc. Adult males are free-swimming. They are usually found in deep waters.

Mystacocarida, crustacean

Class Ostracoda (seed shrimps, mussel shrimps)

Their bodies are enclosed within a bivalve carapace making them look like tiny clams (see below and left). They are all small ranging in size from 0.25 mm to 8.0 mm. The freshwater species rarely exceed 3 mm, although the largest marine species, Gigantocypris mulleri, can be as long as 30 mm.

There are over 8,000 described species, and they have existed since the Cambrian. They are widespread and found in both marine and freshwater habitats at all depths. There are a few terrestrial species found in the moist leaf litter of forest floors.

They have diverse feeding habits including particle, plant, carrion and some are even predators on smaller animals. Some species are luminescent, emitting bluish light in bursts that last 1 - 2 seconds. They are difficult to identify down to species level.When disturbed, the shell valves close tightly, leaving nothing on the outside. From the side they resemble a bean, and from above they look like an egg. Like water fleas the movement of the antennae allow the animal to swim. A plate attached to the jaw vibrates setting up a current of water bringing oxygen to the shell.

The female lays her eggs on water plants. To the naked eye the batch of eggs looks like rust. In favourable conditions they will hatch in 6 - 8 weeks, but they can survive in dry mud as eggs for over 20 years before hatching. On the left is Pionocypris vidua, a freshwater species.

ostracod, seed shrimp, mussel shrimp
Pionocypris vidua, seed shrimp, Ostracoda

Cyclops strenuus
above a female copepod with bunches of eggs, below a parasitic copepod

Ergasilus versicolor, parasitic copepodMacrocyclops albidus, cyclops

Class Copepoda

There are almost 12,000 species described in this Class so far, about 25% are parasitic, and nearly all are less than 2 cm in length. There are around 40 species of cyclops in the UK, all are very similar.

They are found in all kinds of freshwater habitats, where they eat food particles suspended in the water and dead animals. They are mainly marine, but perhaps the best known group are the Cyclops found in freshwater.

They have a characteristic body - club shaped body ending in a forked tail. Usually they have a single black or red eye. They have no carapace or compound eyes. The first pair of antennae is long and the second pair is short.

The non-parasitic Copepods are the dominant members of marine plankton, so are important in the food chain. Left is Cyclops strenuous.

Most non-parasitic species have a life span of one year or less. Some of the freshwater species can secrete a cyst-like covering to help them withstand periods of drying out.

During mating the males use their antennae to grab females. Then the male simply attaches a sperm packet to the female's genitals. Both sexes use their antennae for swimming. Females can have one or two egg sacs each containing around 50 eggs (see left and right). One mating is all that is required to fertilise a few successive set of egg sacs.

Left is Ergasilus versicolor, a parasitic copepod which lives in the gills of freshwater fish. Only the female is parasitic. Note how the second pair of antennae are modified for grasping.

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Class Remipedia

These are small, have an elongated body with about 30 pairs of legs (see left). They are mainly predatory, have many primitive features, and are found in marine caves. !2 species have been described so far.

Class Cephalocardia horseshoe shrimp

There are nine species in this class. All are marine, shrimp-like (see right) and less than 4 mm long. Most have an eight-segmented thorax, and a twelve-segmented abdomen. They have very primitive features, are hermaphroditic, and are without eyes or carapace. They are found in fine sediments.


cepalocardia, crustacean
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